SubstratesMost mangrove species do best on muddy soils, i.e. in areas where siltaccumulates (Watson, 1928; van Steenis, 1957; Chapman, 1976a, 1977;Aragones et al., 1998), and typical for muddy substrates in Southeast Asiaare the well-developed stands of Rhizophora mucronata and Avicennia marina(Watson, 1928; Kint, 1934; van Steenis, 1958). Tall stands of Bruguieradominated forests are often found on deep muddy soils.
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